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Social influences on fertility in group-living mammals

Funding
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Year round
Subject area
Biological and Biomedical Sciences
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Overview

Social competition for limited resources among group-living animals can lead to inhibition of fertility or reproductive suppression, particularly in less competitive individuals that may experience heightened levels of stress. This project will investigate the conditions under which social relationships inhibit or promote the fertility of low-ranking individuals in group-living mammals, with consequences for understanding variation in reproductive skew within and across species. In addition to advancing fundamental understanding of social systems, this project has practical applications in the management of animal populations for conservation breeding.

About this opportunity

Social competition for limited resources among group-living animals can lead to inhibition of fertility or reproductive suppression, particularly in less competitive individuals that may experience heightened levels of stress. However, social relationships can also have beneficial fitness consequences, for example by buffering stress in competitive environments. This project will investigate the conditions under which social relationships inhibit or promote the fertility of low-ranking individuals in group-living mammals, with consequences for understanding variation in reproductive skew within and across species.  In addition to advancing fundamental understanding of social systems, this project has practical applications in the management of animal populations for conservation breeding.

 

Objectives

To investigate how social relationships influence fertility under varying circumstances, the student will design experiments using wild house mice under carefully controlled naturalistic conditions. This approach will allow manipulation of key variables in the social environment (e.g. opportunities for dispersal, levels of within-group relatedness, age asymmetry, resource competition or social group size). Tests will quantify behavioural and physiological responses to contrasting conditions, to assess how changes in the social environment can impact fertility and reproductive skew of group-living mammals.

 

We will also systematically search the literature for published studies relevant to this question, and use a meta-analysis approach to explore broader relationships between behaviour and fertility across group-living mammals with stable social groups.

Examples of key questions to explore include:

  • How do changes to the social environment, for example resulting from delayed dispersal or changes in local relatedness, impact hormonal profiles and fertility of subjects?
  • How do social behaviours, such as the propensity to form social bonds, mediate impacts of social stress on hormonal profiles and fertility?
  • Do females with stronger social relationships tend to have higher reproductive success?
  • How does the number of competitors or social status affect corticosterone levels?

 

This project offers training in a broad range of areas, including evolutionary theory underlying the evolution of social behaviour, the design and implementation of rigorous controlled behavioural

assays, use of R for data manipulation and statistical analysis, approaches for systematic literature review and meta-analysis, and analytical techniques for assaying physiological responses. You will join a productive research environment in the Mammalian Behaviour & Evolution Group at the University of Liverpool’s Leahurst Campus, with outstanding facilities for behavioural research.

Please note that applications may be closed once a suitable candidate is found.

 

Funding notes

This opportunity is for students with their own funding. Funding should cover course fees, living expenses and research expenses (bench fees). The research group cannot provide supplementary funding or provide advice about how to apply for funding. Further information can be found here.

Who is this opportunity for?

Candidates should have a degree in a relevant subject area. Ideally we are looking for candidates with a minimum 2.1 (or equivalent) in their first degree, but we are willing to consider cases where applicants can demonstrate excellence in alternative ways (e.g. performance in masters courses or employment).

Further reading

Fischer, S., Duffield, D., Swaney, W.T., Davidson, A.J., Bolton, R.L., Hurst, J.L. & Stockley, P. (2024) Egalitarian cooperation linked to central oxytocin levels in communally breeding house mice. Communications Biology 7: 1193

 

Fischer, S., Duffield, D., Davidson, A.J., Bolton, R., Hurst, J.L. & Stockley, P. (2023) Fitness costs of female competition linked to resource defence and relatedness of competitors. The American Naturalist. 201, 256-268.

 

Moore, F.R., Shuker, D.M. & Dougherty (2016) Stress and sexual signaling: a systematic review and meta-analysis. Behavioral Ecology 27, 363-371.

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How to apply

  1. 1. Contact supervisors

    Prof Paula Stockley p.stockley@liv.ac.uk https://www.liverpool.ac.uk/people/paula-stockley
    Dr Liam Dougherty Liam.Dougherty@liverpool.ac.uk https://www.liverpool.ac.uk/people/liam-dougherty
    Prof Jane Hurst jane.hurst@liverpool.ac.uk https://www.liverpool.ac.uk/people/jane-hurst

    To apply to this project, in the first instance, please email your CV and cover letter to the primary supervisor: p.stockley@liverpool.ac.uk

  2. 2. Prepare your application documents

    You may need the following documents to complete your online application:

    • A research proposal (this should cover the research you’d like to undertake)
    • University transcripts and degree certificates to date
    • Passport details (international applicants only)
    • English language certificates (international applicants only)
    • A personal statement
    • A curriculum vitae (CV)
    • Contact details for two proposed supervisors
    • Names and contact details of two referees.
  3. 3. Apply

    Finally, register and apply online. You'll receive an email acknowledgment once you've submitted your application. We'll be in touch with further details about what happens next.

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Fees and funding

Your tuition fees, funding your studies, and other costs to consider.

Tuition fees

UK fees (applies to Channel Islands, Isle of Man and Republic of Ireland)

Full-time place, per year - £5,006
Part-time place, per year - £2,505

International fees

Full-time place, per year - £31,249
Part-time place, per year - £15,650

Fees applicable for 2025/26


Additional costs

We understand that budgeting for your time at university is important, and we want to make sure you understand any costs that are not covered by your tuition fee. This could include buying a laptop, books, or stationery.

Find out more about the additional study costs that may apply to this project, as well as general student living costs.

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Contact us

Have a question about this research opportunity or studying a PhD with us? Please get in touch with us, using the contact details below, and we’ll be happy to assist you.

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