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MRC DiMeN Doctoral Training Partnership: Uncovering the molecular mechanisms underpinning A. baumannii virulence

Funding
Funded
Study mode
Full-time
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Start date
Subject area
Biological and Biomedical Sciences

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Overview

Antimicrobial resistance (AMR) is a major global health threat, with Acinetobacter baumannii among the most concerning multidrug-resistant pathogens. This opportunistic bacterium causes severe hospital-acquired infections and is listed by the WHO as a “critical priority” pathogen1. With treatment options increasingly limited, new strategies to combat infection are urgently needed.

About this opportunity

This project takes an alternative approach by targeting bacterial virulence rather than survival, aiming to disarm the pathogen without promoting antibiotic resistance2. The focus is on carboxy-terminal processing proteases (CTPs), a conserved family of serine proteases that regulate cell envelope integrity, signalling, and virulence across all domains of life. Intriguingly, A. baumannii possesses an uncharacterised CTP, which is linked to virulence3, but its molecular function remains unknown.

Research Aims:

This PhD project will uncover how CTPs contribute to A. baumannii virulence and explore their potential as novel therapeutic targets4. The student will define the functional role of the A. baumannii CTP by identifying its interacting proteins and substrates through advanced proteomic approaches, and by evaluating CTP mutants in infection models. Mechanistic studies will focus on how proteolysis is regulated and what determines substrate selectivity, by employing biochemical, biophysical, and structural biology techniques. Peptide-based inhibitors will then be developed and assessed as mechanistic probes to define CTP function and evaluate their impact on pathogen fitness and virulence.

Techniques & Training:

The successful candidate will join an ambitious and supportive research group, dedicated to uncover the molecular mechanisms that underpin bacterial virulence and drive the development of anti-virulence strategies. The project offers exceptional multidisciplinary training across molecular microbiology, infection biology, and structural biochemistry, providing a versatile skillset highly valued in both academia and biotechnology. You will be based in the Department of Clinical Infection, Microbiology & Immunology at the University of Liverpool, benefiting from access to outstanding facilities, technical support, and a vibrant scientific community.

You will receive advanced, hands-on training in cutting-edge methodologies, including:

  • Molecular biology: Core experience in cloning, recombinant protein expression, and purification, forming the foundation for downstream structural and functional studies.
  • Structural biology: Application of X-ray crystallography to resolve the enzyme structures and ligand-bound states, advancing understanding of molecular mechanisms and guiding inhibitor design.
  • Biochemistry & biophysics: Enzyme kinetics, site-directed mutagenesis, and biophysical interaction assays to dissect enzyme activity, regulation, and substrate specificity.
  • Infection biology: Microbial assays and clinically relevant in vivo infection models to assess the physiological role of CTPs and their contribution to virulence.
  • Proteomics & bioinformatics: Quantitative and targeted proteomic approaches to identify enzyme substrates and map protein–protein interaction networks.

Benefits of being in the DiMeN DTP:

This project is part of the Discovery Medicine North Doctoral Training Partnership (DiMeN DTP), a diverse community of PhD students across the North of England researching the major health problems facing the world today. Our partner institutions (Universities of Leeds, Liverpool, Newcastle, York and Sheffield) are internationally recognised as centres of research excellence and can offer you access to state-of-the-art facilities to deliver high impact research.

We are very proud of our student-centred ethos and committed to supporting you throughout your PhD. As part of the DTP, we offer bespoke training in key skills sought after in early career researchers, as well as opportunities to broaden your career horizons in a range of non-academic sectors.

Being funded by the MRC means you can access additional funding for research placements, training opportunities or internships in science policy, science communication and beyond.

Further information on the programme and instructions on how to apply, including a link to the application portal, can be found on our website https://www.dimen.org.uk/

Further reading

1. Morris, F. C., Dexter, C., Kostoulias, X., Uddin, M. I. & Peleg, A. Y. The Mechanisms of Disease Caused by Acinetobacter baumannii. Frontiers in Microbiology 10, doi:10.3389/fmicb.2019.01601 (2019).
2. Lau, W. Y. V., Taylor, P. K., Brinkman, F. S. L. & Lee, A. H. Y. Pathogen-associated gene discovery workflows for novel antivirulence therapeutic development. EBioMedicine 88, 104429, doi:10.1016/j.ebiom.2022.104429 (2023).
3. Roy, R., You, R.-I., Lin, M.-D. & Lin, N.-T. Mutation of the Carboxy-Terminal Processing Protease in Acinetobacter baumannii Affects Motility, Leads to Loss of Membrane Integrity, and Reduces Virulence. Pathogens 9, 322, doi:10.3390/pathogens9050322 (2020).
4. Harding, C. J., Bischoff, M., Bergkessel, M. & Czekster, C. M. An anti-biofilm cyclic peptide targets a secreted aminopeptidase from P. aeruginosa. Nature Chemical Biology 19, 1158-1166, doi:10.1038/s41589-023-01373-8 (2023).

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Who is this for?

This project is ideally suited to a motivated, curious and determined student with a background in biochemistry, molecular biology, or microbiology, who is keen to explore the structural basis of bacterial virulence and contribute to the global fight against antimicrobial resistance.

Applicants for postgraduate research study at Liverpool are normally expected to hold a UK first degree with a First Class or Upper Second Class degree classification, or a Second Class degree plus a Master’s degree. Equivalent international qualifications are also accepted, and their equivalence will be evaluated on the basis of the information provided by the European Network of Information Centres (ENIC) formerly NARIC as well as internal guidance based on our experience of a qualification’s suitability as a preparation for our programmes.

For applicants whose first language is not English, an IELTS score of 6.5 with no band score lower than 5.5, or an equivalent University of Liverpool acceptable English language qualification. For further details and other acceptable English language qualifications please see here: http://www.liv.ac.uk/study/international/countries/english-language/

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How to apply

  1. 1. Contact supervisors

    For informal enquiries or further details, please contact Dr Christopher Harding ().

    More information:

  2. 2. Prepare your application documents

    All applications are made via the application form accessed on the DiMeN website at www.dimen.org.uk

    Please read the full application guidance on the website before submitting an application.

  3. 3. Apply

    Finally, register and apply online. You'll receive an email acknowledgment once you've submitted your application. We'll be in touch with further details about what happens next.

    You should only follow this step if you’ve successfully completed the DiMeN application process

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Funding your PhD

Studentships are fully funded by the Medical Research Council (MRC) for 4yrs. Funding will cover tuition fees, stipend (£20,780 for 2024/25) and project costs. We have a very small number of funded studentships for exceptional international applicants. Please read additional guidance here: View Website

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Contact us

Have a question about this research opportunity or studying a PhD with us? Please get in touch with us, using the contact details below, and we’ll be happy to assist you.

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