Skip to main content
What types of page to search?

Alternatively use our A-Z index.

(BBSRC NWD CASE) Selective breeding vs natural selection: how does body shape influence locomotor biomechanics in domesticated dogs and other mammals?

Funding
Funded
Study mode
Full-time
Apply by
Start date
Subject area
Biological and Biomedical Sciences
Change country or region

We’re currently showing entry requirements and other information for applicants with qualifications from United Kingdom.

Please select from our list of commonly chosen countries below or choose your own.

If your country or region isn’t listed here, please contact us with any questions about studying with us.

Overview

Selective breeding by humans has made the domestic dog the most morphologically diverse species that has ever existed. Since Victorian times, dogs have been the subject of increasingly intense selective breeding with the goal of altering their functional anatomy for specific purposes, such as working tasks, agility or simply ‘desirable’ visual qualities.

About this opportunity

The clear breed-specific prevalence for certain disorders strongly suggests that many of the physical traits favoured by selective breeding have a harmful effect on musculoskeletal function by creating inherent predispositions to disease or injury. Unfortunately, the scientific data needed to understand the relationships between different breed morphologies and locomotor mechanics is currently lacking, limiting the ability of those responsible for canine welfare policy to address health issues surrounding breed standards.

We are attempting to address these issues through quantitative comparisons of anatomy and gait dynamics in popular UK domestic dog breeds. At present, however, we have not assessed the impacts of differences in anatomy and gait on the loading of joint tissues (ligament, cartilage and bone) across breeds, which could be carried out non-invasively using finite element analysis. Furthermore, our work is yet to place the anatomical and movement variation seen in dogs in a wider context by comparing to other selectively bred mammals or indeed animals that evolved purely by natural selection. Such an analyses could have major welfare impacts, as well as allowing us to use body shape variation seen in selectively bred animals as model systems to understand how anatomical variation alters quadrupedal locomotor mechanics.

The goal of this project is to determine if differences in anatomy and habitual motion patterns across canine breeds result in joint tissue loads and safety factors that different not only to each other but also a range of other domesticated and non-domesticated mammals, through the following objectives:

  1. To build and analyse breed-specific finite element computer models of knee and elbow joints and load these models with existing kinetic data from experimental gait analysis and MDA models.
  2. Conduct comparative morphological, experimental and modelling work to quantitatively compare the magnitude of anatomical change and the resulting variation in bone loading seen in domestic dog breeds to ancestral and wild dogs, and other selectively bred or domesticated mammals (e.g. cats, horses).
  3. In collaboration with SME Fusion Implants (CASE partner), use our experimental and FE modelling framework to assess a new Load Shifting Humeral Osteotomy device for large-breed dogs.

The ideal student would have a background in zoology, comparative anatomy or bioengineering, and skills in quantitative, mechanical and/or 3D digital techniques, and/or experimental gait analysis, but training will be provided in all techniques. The supervisory team includes experts in mammalian anatomy, biomechanics, computer simulation, tissue mechanics and canine welfare, and some flexibly exists to customise the project to the interest of successful student.

Further reading

1. Charles, J.P., Comerford, E., Ratcliffe, V., Kissane, R.W.P., Gooding, I., Cottriall, S., Maddox, T.W. & Bates, K.T. 2025. Biomechanics of working dog locomotion I: steady-state trotting. Journal of Experimental Biology.
2. Charles, J.P., Comerford, E., Ratcliffe, V., Kissane, R.W.P., Gooding, I., Cottriall, S., Maddox, T.W. & Bates, K.T. 2025. The biomechanics of working dog locomotion II: Loaded trotting. Journal of Experimental Biology.
3. Clark, N.L., Bates, K.T., Murray, J., Harris, L. & Comerford, E. 2023. GenPup-M: a novel validated owner-reported clinical metrology instrument for detecting early mobility changes in dogs. PLoS ONE 18(12): e0291035.
4. Sellers, W.I., Cross, C.F., Fukuhara, A., Ishiguro, A. & Hirasaki, E. 2023. Producing non-steady-state gaits (starting, stopping, and turning) in a biologically realistic quadrupedal simulation. Frontiers in Ecology & Evolution 10, 954838.
5. Readioff, R., Geraghty, B., Comerford, E., & Elsheikh, A. 2020. A Full-Field 3D Digital Image Correlation and Modelling Technique to Characterise Anterior Cruciate Ligament Mechanics Ex Vivo. Acta Biomaterialia 113:417–28.

Back to top

Who is this for?

Applicants must have obtained or be about to obtain a minimum Upper Second class UK honours degree, or the equivalent qualifications gained outside the UK, in an appropriate area of science, engineering or technology.

International applicants

We are only able to offer a limited number of full studentships to applicants outside the UK. Therefore, full studentships will only be awarded to exceptional quality international candidates due to the competitive nature of this scheme.

International applicants must ensure they meet the academic eligibility criteria (including English language) before applying. Visit our English Language requirements page to find out more.

Equality, Diversity and Inclusion

Equality, diversity and inclusion is fundamental to the success of The University of Liverpool, and is at the heart of all of our activities. The full equality, diversity and inclusion statement can be found on our website.

Back to top

How to apply

  1. 1. Contact supervisors

    The student will be based with Professor Bates in the Evolutionary Morphology & Biomechanics Group at Liverpool (https://www.liverpool.ac.uk/life-course-and-medical-sciences/research/groups/evolutionary-morphology-and-biomechanics-group/) but will spend time in the labs of the co-supervisors at Liverpool and Manchester.

  2. 2. Prepare your application documents

    Browse our BBSRC NWD in Bioscience projects and discover one you’re passionate about that matches your interests, ambitions and goals.

    Applicants must make direct contact with preferred supervisors before applying. It is your responsibility to make arrangements to meet with potential supervisors, prior to submitting a formal online application.

    How to Apply

    All applications should be submitted through the University of Manchester application portal.

    Apply directly via this link, and select BBSRC DTP PhD as the programme of study. You may apply for up to two projects from the programme via this scheme. To do so, submit a single online application listing both project titles and the names of both main supervisors in the relevant sections.

    Please ensure that your application includes all required supporting documents:

    • Curriculum Vitae (CV)
    • Supporting Statement
    • Academic Certificates and Transcripts

    Incomplete or late applications will not be considered.

    Applications should not be made through the University of Liverpool’s application portal.

    You must submit your application form along with the required supporting documents by the deadline date. You can select up to two projects on one single application, noting the title of each project from the advert and the supervisor name. This can include two projects from one institution or a project from each institution.

    Once you have completed your application, you’ll receive a confirmation email.

    Deadline: Sunday 7th December, midnight (UK time)

    Late or incomplete applications will not be considered.

    If you need help with this stage of the process, or have any queries regarding your eligibility (such as if you achieved unexpectedly low degree results due to extenuating circumstances), please contact the Liverpool BBSRC team for advice at 

  3. 3. Apply

    Finally, register and apply online. You'll receive an email acknowledgment once you've submitted your application. We'll be in touch with further details about what happens next.

    Once you have applied through the University of Manchester portal, and if you are successfully offered a studentship following a formal interview, you will be instructed to apply formally through the University of Liverpool. You must only do this once you have been instructed to do so.

Back to top

Funding your PhD

This is a 4 year CASE studentship in partnership with Fusion Implants available to UK and international applicants, and provide funding for tuition fees and stipend at the UKRI rate, subject to eligibility, for four years.. This covers tuition fees and an annual stipend. This does not include any costs associated with relocation.

Back to top

Contact us

Have a question about this research opportunity or studying a PhD with us? Please get in touch with us, using the contact details below, and we’ll be happy to assist you.

Back to top